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Currently, contains four subspecies: subsp. , subsp. , subsp. and subsp. . In the study of Pérez , these four subspecies could be clearly divided into two groups based on sequence analysis: subsp. and subsp. ; subsp. and subsp. . The two groups had a relatively low DNA-DNA hybridization value (about 60 %). In the present study, the taxonomic position of subsp. and subsp. was re-examined based on sequence analyses of 16S rRNA, , and genes, average nucleotide identity (ANI) values and digital DNA-DNA hybridization (dDDH) values. The result of 16S rRNA gene sequence analysis indicated that subsp. NCDO 607 and subsp. L105 were phylogenetically related to the type strains of subsp. , subsp. , , , , , , and . The 16S rRNA gene, , , and concatenated , and sequence similarities, ANI values, and dDDH values between the type strains of subsp. , subsp. and phylogenetically related species were 93.5-99.4 %, 83.3-97.6 %, 80.6-92.4 %, 79.7-92.7 %, 83.5-94.3 %, 72.4-86.9 % and 21.4-32.5 %, respectively. Lower than 95-96 % ANI values and lower than 70 % dDDH values confirmed that the type strains of subsp. and subsp. represent a novel species in the genus . Because subsp. was proposed and validated before subsp. , subsp. is elevated to the species level as sp. nov. and subsp. is transferred to as subsp. comb. nov. The type strain of sp. nov. is NCDO 607 (=ATCC 19257=DSM 20069=JCM 16167=LMG 6897=NBRC 100676). The type strain of subsp. comb. nov. is L105 (=NBRC 110453=DSM 21502=JCM 31125=LMG 24662).