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Marine Biological Association of the United Kingdom, The Laboratory, , Citadel Hill, Plymouth PL1 2PB, UK, European Institute of Marine Studies (IUEM), University of Western Britanny, , Place Nicolas Copernic 29280, Plouzané, France, School of Computing and Mathematics, University of Plymouth, , Drake Circus, Plymouth PL4 8AA, UK, Rothamsted Research, , Harpenden AL5 2JQ, UK, Ocean and Earth Science, National Oceanography Centre Southampton, University of Southampton, , Waterfront Campus, Southampton SO14 3ZH, UK, Centre for Biological Sciences, University of Southampton, , Building 85, Highfield Campus, Southampton SO17 1BJ, UK.